Connelly was born in Round Top, New York in the Catskill Mountains to Ilene, an antiques dealer, and Gerard Connelly, a clothing manufacturer.Her father was a Roman Catholic with Irish and Norwegian ancestors,while her mother was a yeshiva-educated Jew of Polish and Russian stock. She was first raised in Brooklyn Heights, near the Brooklyn Bridge, where she attended Saint Ann's, a private school focused on the arts. The family moved to Woodstock, New York in 1976 Four years later the family returned to Brooklyn Heights where Connelly resumed her education at the same school.
After Connelly's frequent appearances as a model, her mother started to take her to acting auditions where she was selected for a supporting role as Deborah Gelly in Sergio Leone's 1984 gangster epic, Once Upon a Time in America. In a scene of the film, Connelly had to perform a ballet routine. During the audition for the role, and without any knowledge of the dance, she made random movements imitating it that finally convinced the director to include her in the cast. Connelly described the movie as "an incredibly idyllic introduction to movie-making". Her first leading role was as Jennifer Corvino in Italian giallo-director Dario Argento's 1985 film Phenomena,followed by the lead in the coming-of-age movie Seven Minutes in Heaven the same year.
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